After the krogan respond to the initial turian offensive by devastating turian colonies with weapons of mass destruction, the turians vow to stop the krogan from ever becoming a threat again.
The Citadel Council makes first contact with the turians around this time and persuades them to aid in the war. A preemptive strike is made on krogan infrastructures by the Spectres. 
Eventually the Council demands withdrawal from the asari colony of Lusia, but the krogan refuse.
Krogan warlords leverage veterans of the Rachni Wars to annex territory from other races in Citadel space. Free of the harsh environment of Tuchanka, the krogan population explodes. In gratitude for their aid during the Rachni Wars, the Council rewards the krogan a new homeworld. The krogan prove able to survive the harsh environments of the rachni worlds and pursue the rachni into their nests, systematically eradicating queens and eggs. 
The salarians make first contact with and uplift the primitive krogan, manipulating them into acting as soldiers for the Citadel Council. Negotiation with the rachni queens is impossible because they cannot be contacted in their underground nests on the toxic rachni worlds. The rachni prove to be hostile and begin a war with the rest of the galaxy.
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The rachni, a species of highly intelligent hive-minded insects, are discovered when a Citadel expedition opens a dormant mass relay leading to their star systems. The elcor and the quarians are granted embassies on the Citadel. First contact is made with the quarians.
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